Movement also telling story because it gives character individual. That’s the thought came out after watching Winsor McCay’s short animations. Comparing with full-length animation/cartoon, most short animation film cannot build a long personal background about characters so those movements animators give to characters’ usually let audience discover characters’ individual directly and quickly.
I think How a Mosquito Operates (1912) is a really good example about storytelling come from movements.https://youtu.be/lmVra1mW7LU
Characters’ movement even more important than real actors in film, because everyone knows that characters are made by animators, they always come from some reasons. So wrong/ unclear movements will let them misunderstand.
Some reference from Richard Williams